15th May 08

Drosophila melanogaster

Drosophila melanogaster is a fruit fly and happens to be one of the most valuable organisms to biological research. It has been used as a model organism for nearly 100 years, particularly in biological areas of genetics and development. It is the second multicellular organism to have its genome completely sequenced following C. elegans.

Many genes found in Drosophila have equivalents in higher organisms, hence study of the Drosophila genome can relay information regarding human genes. Most of the genes that encode spatial pattern of cell types and body parts have now been identified, such as those involved with hedgehog signalling.

The Drosophila genome is ~165 Mb and contains 13,000 genes. This fruit fly has four pairs of chromosomes: the X/Y sex chromosomes and the autosomes 2, 3 and 4. Chromosome 4 is relatively small.

Embryonic Cell Fate:

During animal embryonic development, localized Hh signals organise patterns and cell types by controlling cell fates. In Drosophila, embryonic segments and nascent imaginal discs are divided into anterior (A) and posterior (P) compartment cells. A cells transcribe Ci, a transcription factor and P cells transcribe engrailed (en), a homeobox containing transcription factor.

A Hh signal from P cells, allows transcription of wg signalling molecules in embryonic development and dpp in leg and wing imaginal discs.

When A cells receive a Hh signal, Ci directly activates Hh target gene transcription. This is thought to be allowed by an increase in Fu activity and decrease in Cos2 activity.

Overview
C. Elegans
Signalling
Hedgehog
Signalling
Pathway
Components
C. Elegans
C. Briggsae
Drosophila
Homologues
Targets
Database
 
 
 
Links:

The University
of Liverpool


C elegans Blast

Sanger Institute

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